中华锹甲 2PDF电子书下载

Furthur research on the genera Lucanus Scopoli and Eolucanus Kurosawa from China,with the description of two new species 1

An updated checklist of Chinese species of the genera Lucanus and Eolucanus 1

Notes on some species of the genera Lucanus and Eolucanus 3

Ⅰ．Lucanus wuyishanensis Schenk武夷深山 3

Ⅱ．Lucanus liuyei Huang ＆ Chen刘氏深山 4

Ⅲ．Lucanus swinhoei Parry姬深山 5

Ⅳ．Lucanus prossi Zilioli普氏深山 5

Ⅴ．Lucanus brivioi Zilioli布氏深山 6

Ⅵ．Lucanus nobilis Didier雅深山 6

Ⅶ．Lucanus dirki Schenk大理深山 6

Ⅷ．Lucanus formosus Didier橙深山 7

Ⅸ．Lucanus fujitai Katsura ＆ Giang北越深山 7

Ⅹ．Lucanus sp．incert．墨脱未定名深山 8

Ⅺ．Lucanus delavayi Fairmaire黄鞘深山 8

Ⅻ．Lucanus parryi parryi Boileau帕瑞深山指名亚种 9

ⅩⅢ．Lucanus fujianensis Schenk微齿姬深山 9

ⅩⅣ．Lucanus fairmairei Planet四川深山 10

ⅩⅤ．Lucanus laminifer Waterhouse拉叉深山 10

ⅩⅥ．Lucanus angusticornis inclinatus Schenk弯叉深山中国亚种 11

ⅩⅦ．Lucanus hermani De Lisle巨叉深山 11

ⅩⅧ．Lucanus planeti planeti Planet普叉深山指名亚种 11

ⅩⅨ．Lucanus planeti dayaoshanensis Schenk普叉深山广西亚种 11

ⅩⅩ．Lucanus dybowski lhasaensis Schenk斑股深山四川亚种 12

ⅩⅪ．Lucanus ludivinae Boucher路氏深山 13

ⅩⅫ．Lucanus lunifer Westwood藏南深山 14

ⅩⅩⅢ．Lucanus thibetanus furcifer Arrow藏深山云南亚种 15

ⅩⅩⅣ．Lucanus thibetanus singularis Planet藏深山怒江亚种 16

ⅩⅩⅤ．Lucanus thibetanus isaki Nagai藏深山独龙江亚种 22

ⅩⅩⅥ．Lucanus thibetanus ssp藏深山片马未定亚种 23

ⅩⅩⅦ．Lucanus langi Huang,He ＆ Shi郎氏深山 23

ⅩⅩⅧ．Lucanus fryi Boileau弗瑞深山 25

ⅩⅩⅨ．Lucanus atratus Hope阿深山 25

ⅩⅩⅩ．Lucanus mearesii Hope金属深山 26

ⅩⅩⅪ．Lucanus nyshiwini bretschneideri Schenk双齿深山墨脱亚种 26

ⅩⅩⅫ．Eolucanus gracilis（Albers）短颚拟深山 27

ⅩⅩⅩⅢ．Eolucanus lesnei（Planet）尖齿拟深山 27

ⅩⅩⅩⅣ．Eolucanus mingyiae（Huang）圆齿拟深山 27

ⅩⅩⅩⅤ．Eolucanus pani（Huang）潘氏拟深山 30

ⅩⅩⅩⅥ．Eolucanus oberthuri（Planet）欧氏拟深山 30

Further notes 30

New descriptions 30

ⅩⅩⅩⅦ．Lucanus choui Huang ＆ Chen sp．nov．周氏深山（新种） 30

ⅩⅩⅩⅧ．Lucanus hewenjiae Huang ＆ Chen sp．nov．何氏深山（新种） 33

Descriptions of some new taxa of Lucanidae from China 35

A．Prismognathus zhangi Huang ＆ Chen sp．nov．张氏鬼锹（新种） 35

B．Ceruchus yingqii Huang ＆ Chen sp．nov．浅纹铠锹（新种） 36

C．Himaloaesalus Huang ＆ Chen gen．nov．喜马拉雅斑锹属（新属） 39

Phylogeny and systematics of Lucaninae （sensu Holloway,1960 ＆ 1968）from China 41

Introduction 41

Abbreviations 41

Choice of ingroup and outgroup taxa 41

Source of data and initial names 43

Morphology 43

Characters coding and data sets 45

Softwares 46

Analyses and searching results 47

Monophyla and higher taxa 62

1） Genera 62

2） Tribes 64

3） Species groups in Lucanus 65

4） Species groups in Neolucanus 66

5） Species groups in Prosopocoilus,Dorcus and Falcicornis 66

6） Clade of Sinodendron 66

Systematics and annotated list of higher taxa of Chinese Lucanidae,with some extension to species from outside China,and with emphasis on the tribe Dorcini 66

Unsorted genera from China 66

Subfamily AESALINAE MacLeay,1819 67

Tribe Aesalini MacLeay,1819 67

Subfamily SYNDESINAE MacLeay,1819 67

Tribe Sinodendrini Mulsant,1842 67

Tribe Ceruchini Jacquelin du Val ＆ Fainnaire,1859 67

Subfamily LUCANINAE Latreille,1804 67

Tribe Platycerini Mulsant,1842 67

Tribe Noseolucanini Huang ＆ Chen,2013,new tribe 67

Tribe Aegini Huang ＆ Chen,2013,new tribe 68

Tribe Odontolabini Parry,1870 69

Tribe Figulini Burmeister,1847 69

Tribe Capreolucanini Huang ＆ Chen,2013,new tribe 69

Tribe Cyclommatini Huang ＆ Chen,2013,new tribe 70

Tribe Lucanini Latreille,1804 71

Tribe Dorcini Parry,1864 72

Genus Gnaphaloryx Burmeister,1847 72

Genus Kirchnerius Schenk,2009 72

Genus Metallactulus Ritsema,1885 73

Genus Epidorcus Seguy,1954 73

Genus Serrognathus Motschulsky,1861 74

Genus Hexarthrius Hope,1842 75

Genus Rhaetus Parry,1864 75

Genus Rhaetulus Westwood,1871 76

Genus Pseudorhaetus Planet,1899 76

Genus Prosopocoilus Hope ＆ Westwood,1845 76

Genus Miwanus Huang ＆ Chen,new genus 77

Genus Falcicornis Planet,1894 78

Genus Sinodorcus Huang ＆ Chen,new genus 78

Genus Aulacostethus Waterhouse,1869 79

Genus Dorcus MacLeay,1819 79

Discussion on some tribes from outside China 80

Key to tribes of Lucaninae from China 82

Appendix 1-List of characters used for phylogenetic analyses 84

Appendix 2-Data matrix 101

A review of Dorcini（sensu Huang ＆ Chen,2013）from China 115

Introduction 115

Key to genera of Dorcini males 115

Key to genera of Dorcini females 117

Genus Gnaphaloryx Burmeister,1847黑锈锹属 119

1．Gnaphaloryx opacus Burmeister,1847黑锈锹 119

Genus Kirchnerius Schenk,2009钳锹属 121

Key to Chinese species of Kirchnerius males 121

Key to Chinese species of Kirchnerius females 122

Kirchnerius spencei group 122

2．Kirchnerius spencei（Hope,1840）comb．nov．钳锹 122

Kirchnerius boreli group 125

3．Kirchnerius guangxii Schenk,2009 广西钳锹 125

4．Kirchnerius yangi（Fukinuki,2004）comb．nov．杨氏钳锹 127

5．Kirchnerius simianshanus（Huang ＆ Chen,2011）comb．nov．四面山钳锹 128

6．Kirchnerius boreli（Boileau,1904）comb．nov．印度钳锹 129

Genus Metallactulus Ritsema,1885姬扁锹属 131

7．Metallactulus parvulus（Hope,1845）姬扁锹 131

Genus Epidorcus Séguy,1954前锹属 132

Key to Chinese species of Epidorcus males 132

Key to Chinese species of Epidorcus females 133

8．Epidorcus gracilis（Saunders,1854）狭长前锹 134

9．Epidorcus piceipennis（Westwood,1855）拟狭长前锹 137

10．Epidorcus denticulatus（Boileau,1901）,comb．nov．单基齿前锹 139

11．Epidorcus rondoni（Bomans ＆ Lacroix,1970）,comb．nov．短基齿前锹 144

12．Epidorcus tonkinensis（Pouillaude,1913）,comb．nov．并基齿前锹 145

13．Epidorcus bidentatus（Bomans,1978）,comb．nov．双基齿前锹 147

Genus Serrognathus Motschulsky,1861扁锹属 147

Key to Chinese taxa of Serrognathus males 147

Key to Chinese taxa of Serrognathus females 148

14．Serrognathus titanus（Boisduval,1835）扁锹 149

Serrognathus titanus platymelus（Saunders,1854）扁锹典型亚种 149

Serrognathus titanus fafner（Kriesche,［1921］）扁锹越南亚种 154

Serrognathus titanus typhoniformis（Nagel,1924）扁锹云贵亚种 155

Serrognathus titanus sika（Kriesche,［1921］）扁锹台湾亚种 156

Serrognathus titanus castanicolor Motschulsky,1861扁锹东北亚种 157

Serrognathus titanus ssp．incert．1扁锹版纳亚种 159

Serrognathus titanus ssp．incert．2扁锹墨脱亚种 160

Species delimitation 161

Diagnosis for species 163

15．Serrognathus consentaneus（Albers,1886）尖腹扁锹（别名：细齿扁） 165

16．Serrognathus kyanrauensis（Miwa,1934）深山扁锹 168

Genus Hexarthrius Hope,1842六节锹属 169

Key to Chinese taxa of Hexarthrius males 169

Key to Chinese taxa of Hexarthrius females 170

17．Hexarthrius aduncus Jordan,1894印度六节锹 170

18．Hexarthrius vitalisi Didier,1925维氏六节锹 176

Hexarthrius vitalisi vitalisi Didier,1925维氏六节锹指名亚种 176

Hexarthrius vitalisi tsukamotoi Nagai,1998维氏六节锹桂北亚种 177

Hexarthrius vitalisi ssp．incert．1维氏六节锹桂西亚种 178

Hexarthrius vitalisi ssp．incert．2维氏六节锹高黎贡山亚种 178

Species delimitation 179

Subspecies 179

19．Hexarthrius partyi Hope,1842橘背六节锹（别名：橘背叉角、帕瑞六节） 179

20．Hexarthrius forsteri（Hope,1840）佛氏六节锹 182

21．Hexarthrius melchioritis Séguy,1954缅甸六节锹 183

Genus Rhaetus Parry,1864巨鹿锹属 185

22．Rhaetus westwoodi（Parry,1862）巨鹿锹（别名：金刚鹿） 185

Genus Rhaetulus Westwood,1871鹿角锹属 187

23．Rhaetulus crenatus Westwood,1871鹿角锹 187

Rhaetulus crenatus crenatus Westwood,1871鹿角锹指名亚种 187

Rhaetulus cretatus rubrifemoratus Nagai,2000鹿角锹华南亚种 189

Rhaetulus crenatus boileaui Didier,1925？鹿角锹版纳亚种 190

Rhaetulus crenatus ssp．incert．鹿角锹中缅亚种 191

Species delimitation 191

Subspecies 191

Genus Pseudorhaetus Planet,1899拟鹿锹属 192

Key to Chinese taxa of Pseudorhaetus 192

24．Pseudorhaetus sinicus（Boileau,1899）拟鹿锹（别名：漆黑鹿角） 192

Pseudorhaetus sinicus sinicus（Boileau,1899）拟鹿锹指名亚种 192

Pseudorhaetus sinicus concolor Benesh,1960拟鹿锹台湾亚种 193

Species delimitation 194

Diagnosis for species 194

25．Pseudorhaetus oberthuri Planet,1899红背拟鹿锹 194

Genus Prosopocoilus Hope,1845锯锹属 195

Key to Chinese species of Prosopocoilus males 195

Key to Chinese species of Prosopocoilus females 197

Prosopocoilus buddha group 199

26．Prosopocoilus approximatus（Parry,1864）伪布达锯锹 199

27．Prosopocoilus thibeticus（Westwood,1855）滇东南锯锹 203

28．Prosopocoilus porrectus Bomans,1978歧齿锯锹 206

29．Prosopocoilus fulgens（Didier,1927）毛刷锯锹 206

30．Prosopocoilus confucius（Hope,1842）孔夫子锯锹 208

31．Prosopocoilus giraffa giraffa（Olivier,1789）长颈鹿锯锹指名亚种 209

32．Prosopocoilus politus（Parry,1862）亮锯锹 210

33．Prosopocoilus forficula（Thomson,1856）圆翅锯锹 212

Prosopocoilus astacoides group 214

34．Prosopocoilus astacoides（Hope,1840）两点锯锹 214

Species delimitation and subspecies delimitation 214

Prosopocoilus astacoides astacoides（Hope,1840）两点锯锹指名亚种 217

Prosopocoilus astacoides castaneus（Hope,1845）两点锯锹樟木亚种 219

Prosopocoilus astacoides fraternus（Hope,1845）两点锯锹滇南亚种 221

Prosopocoilus astacoides kachinensis Bomans ＆ Miyashita,1997两点锯锹藏东南亚种 223

Prosopocoilus astacoides blanchardi（Parry,1873）两点锯锹普通亚种 224

Prosopocoilus astacoides dubernardi（Planet,1899）两点锯锹滇西北亚种 226

Other subspecies 227

Diagnosis for species 227

Prosopocoilus impressus（Waterhouse,1869）-Valid species,not found in China 229

Prosopocoilus laterinus（Didier,1928）-Not found in China 230

35．Prosopocoilus reni Huang ＆ Chen,2011任氏锯锹 231

36．Prosopocoilus yazakii Nagai,2005三色锯锹 231

Prosopocoilus inclinatus group 232

37．Prosopocoilus inclinatus inclinatus（Motschulsky,［1858］）东北锯锹 232

38．Prosopocoilus motschulskyi（Waterhouse,1869）高砂锯锹 234

39．Prosopocoilus oweni melli Kriesche,1922欧文锯锹中国亚种 236

40．Prosopocoilus doris Kriesche,［1921］平齿锯锹 238

41．Prosopocoilus suturalis（Olivier,1789）丫纹锯锹 240

42．Prosopocoilus biplagiatus（Westwood,1855）黄纹锯锹 241

43．Prosopocoilus spineus（Didier,1927）红背锯锹 243

Prosopocoilus maclellandi group 245

44．Prosopocoilus jenkinsi（Westwood 1848）毛胫四点锯锹 245

45．Prosopocoilus maclellandi（Hope,1842）四点锯锹 249

46．Prosopocoilus laterotarsus（Houlbert,1915）砂缘锯锹 251

Genus Miwanus Huang ＆ Chen,2013 gen．nov．双钩锹属（新属） 254

47．Miwanus formosanus（Miwa,1929）comb．nov．双钩锹 254

Miwanus formosanus formosanus（Miwa,1929）双钩锹指名亚种（台湾亚种） 254

Miwanus formosanus capricornus（Didier,1931）双钩锹越南亚种 255

Miwanus formosanus kishidai（Fujita,2010）双钩锹广东亚种 256

Miwanus formosanus ssp．incert．双钩锹华中未定名亚种 256

Species delimitation 257

Genus Falcicornis Planet,1894小刀锹属 257

Key to Chinese species of Falcicornis males 257

Key to Chinese species of Falcicornis females 259

48．Falcicornis neolucanoides sp．nov．圆翅小刀锹（新种） 261

Falcicornis elegantulus group 262

49．Falcicornis pseudaxis（Didier,1926）comb．nov．南红小刀锹 262

50．Falcicornis mellianus（Kriesche,［1921］）comb．nov．黄毛小刀锹 264

51．Falcicornis seguyi（De Lisle,1955）comb．nov．叉齿小刀锹 266

Falcicornis vernicatus group 269

52．Falcicornis humilis（Arrow 1935）comb．nov．西藏小刀锹 269

53．Falcicornis vernicatus（Arrow,1938）comb．nov．戟小刀锹 276

54．Falcicornis itoi（Bomans,1993）comb．nov．越南小刀锹 279

55．Falcicornis taibaishanensis（Schenk,2008）comb．nov．拟戟小刀锹 279

56．Falcicornis heyangi Huang ＆ Chen sp．nov．何洋小刀锹（新种） 280

Falcicornis bisignatus group 281

57．Falcicornis tenuecostatus（Fairmaire,1888）comb．nov．皮氏小刀锹 281

Falcicornis tenuecostatus tenuecostatus（Fairmaire,1888）皮氏小刀锹华北亚种 281

Falcicornis tenuecostatus pieli（Didier ＆ Séguy,1953）stat．nov．皮氏小刀锹华南亚种 283

Falcicornis tenuecostatus mochizukii（Miwa,1937）stat．nov．皮氏小刀锹台湾亚种（望月锹） 286

Falcicornis tenuecostatus ssp．incert．皮氏小刀锹西康亚种 286

Species delimitation and diagnosis for species 287

58．Falcicornis moellenkampi（Nagel,1924）comb．nov．莫氏小刀锹 290

Falcicornis moellenkampi moellenkampi（Nagel,1924）莫氏小刀锹指名亚种 290

Falcicornis moellenkampi bomansi（Lacroix,1981）stat．nov．莫氏小刀锹滇西亚种 291

Diagnosis for species 293

Records 294

59．Falcicornis ruficrus（De Lisle,1970）红河小刀锹 294

60．Falcicornis bisignatus（Parry,1862）束胸小刀锹 295

61．Falcicornis fulvonotatus（Parry,1862）斑胸小刀锹 300

62．Falcicornis rufonotatus （Pouillaude,1913）滇越小刀锹 302

63．Falcicornis songianus（Didier ＆ Séguy,1953）华南小刀锹 304

64．Falcicornis himalayae（Schenk 2009）宽胸小刀锹 307

Genus Sinodorcus Huang ＆ Chen,2013 gen．nov．多刺刀锹属（新属） 308

65．Sinodorcus sawaii（Tsukawaki,1999）comb．nov．多刺刀锹 308

Genus Aulacostethus Waterhouse,1869阿锹属 309

Key to species of Aulacostethus 309

66．Aulacostethus archeri Waterhouse,1869印度阿锹 310

67．Aulacostethus ruditemporalis（Houlbert,1914）四川阿锹 311

68．Aulacostethus doani Baba,2000越南阿锹 312

69．Aulacostethus tianmuxing Huang ＆ Chen sp．nov．华东阿锹（新种） 313

Genus Dorcus MacLeay,1819刀锹属、大锹属 314

Key to Chinese species of Dorcus males 315

Key to Chinese species of Dorcus females 326

Dorcus antaeus group 333

70．Dorcus schenklingi（M？llenkamp,1913）长角大锹 333

71．Dorcus antaeus Hope,1842安达大锹（别名：安达佑实大锹） 335

72．Dorcus vicinus Saunders,1854华东大锹（别名：天章大锹） 339

73．Dorcus rectus（Motschulsky,［1858］）单齿刀锹（别名：日本小锹、直牙锹） 341

Dorcus montivagus group 344

74．Dorcus davidis（Fairmaire,1887）大卫大锹 344

Dorcus rubrofemoratus group 348

75．Dorcusyamadai（MIWA,1937）台湾刀锹（别名：刀锹） 348

76．Dorcus rubrofemoratus（van Vollenhoven,1865）红腿刀锹 351

Dorcus rubrofemoratus rubrofemoratus（van Vollenhoven,1865）红腿刀锹指名亚种 351

Dorcus rubrofemoratus chenpengi（Li,1992）红腿刀锹北部亚种 352

Dorcus rubrofemoratus miyamai Nagai ＆ Okazaki,2005红腿刀锹西部亚种 353

Species delimitation and diagnosis for species 354

77．Dorcus haitschunus（Didier ＆ Séguy,1952）长齿刀锹 354

78．Dorcus arrowi（Boileau,1911）红背刀锹 356

Dorcus arrowi arrowi（Boileau,1911）红背刀锹指名亚种（滇西亚种） 356

Dorcus arrowi magdaleinae（Lacroix,1972）stat．nov．红背刀锹滇南亚种 358

Dorcus arrowi ssp．incert．红背刀锹滇东南亚种 360

Diagnosis for species 362

79．Dorcus derelictus Parry,1862短颚刀锹 363

Dorcus nepalensis group 364

80．Dorcus sinensis（Boileau,1899）中华刀锹 364

81．Dorcus semenowi Jakowlew,1900谢苗诺夫刀锹 371

82．Dorcus kikunoae Hosoguchi,2004错那刀锹 377

83．Dorcus wui Huang ＆ Chen sp．nov．吴氏刀锹（新种） 378

84．Dorcus ratiocinativus Westwood,1871弯颚刀锹 380

85．Dorcus wardi Arrow,1943沃德刀锹 381

86．Dorcus macleayii（Hope,1845）麦氏刀锹 383

87．Dorcus branaungi Nagai,2000布氏刀锹（别名：布朗刀） 386

88．Dorcus sp．incert．1二郎山刀锹 387

89．Dorcus sp．incert．2巨刀锹 388

90．Dorcus nepalensis（Hope,1831）尼泊尔刀锹 389

91．Dorcus donckieri（Boileau,1898）东氏刀锹 392

Dorcus curvidens group 393

92．Dorcus curvidens（Hope,1840）弯角大锹 393

93．Dorcus hopei（Saunders,1854）中华大锹（别名：中国大锹） 397

Dorcus hopei hopei（Saunders,1854）中华大锹指名亚种 397

Dorcus hopei formosanus Miwa,1929 stat．nov．中华大锹台湾亚种 400

Diagnosis for species 401

Subspecies 402

94．Dorcus yaksha Gravely,1915细角大锹 402

Dorcus yaksha yaksha Gravely,1915细角大锹指名亚种 402

Dorcus yaksha gracilicornis Benesh,1950 stat．nov．细角大锹东部亚种 404

Dorcus katsurai group 407

95．Dorcus katsurai Ikeda,2000扩胫大锹 408

96．Dorcus sukkiti Fukinuki,2004缅甸扩胫大锹 411

97．Dorcus sp．incert．3肘子大锹 411

98．Dorcus myinti Nagai ＆ Maeda,2009敏氏刀锹 411

Dorcus velutinus group 413

99．Dorcus velutinus Thomson,1862锈刀锹 413

100．Dorcus tenuihirsutus Kim ＆ Kim,2010北方锈刀锹 418

101．Dorcus ursulus Arrow 1938扩胸锈刀锹 419

102．Dorcus taiwanicus Nakane ＆ Makino,1985台湾锈刀锹 420

103．Docus carinulatus Nagel,1941直颚锈刀锹 422

104．Dorcus ursulae（Schenk,1996）平齿刀锹 424

105．Dorcus tanakai Nagai,2002田中氏刀锹 426

Dorcus elegans group 427

106．Dorcus elegans（Parry,1862）樟木怪刀锹 427

107．Docus motuoensis Huang ＆ Chen sp．nov．墨脱怪刀锹（新种） 429

108．Dorcus chucheni Huang ＆ Chen sp．nov．初氏怪刀锹（新种） 433

109．Dorcus wemckeni（Schenk,2008）短颚怪刀锹 434

Dorcus striatipennis group（sensu stricto） 435

110．Dorcus striatipennis（Motschulsky,1861）双齿刀锹 435

Dorcus striatipennis yushiroi Sakaino,1997双齿刀锹台湾亚种 435

Dorcus striatipennis davidi（Séguy,1954）stat．nov．双齿刀锹大陆亚种 436

Delimitation of species 439

Subspecies 441

111．Dorcus lvbu Huang ＆ Chen sp．nov．吕布刀锹（新种） 442

112．Dorcus bashanus Huang ＆ Chen sp．nov．巴山刀锹（新种） 445

113．Dorcus gongshanus Huang ＆ Chen sp．nov．贡山刀锹（新种） 446

114．Dorcus intricatus（Lacroix,1981）雅安刀锹 448

115．Dorcus linwenhsini Huang ＆ Chen sp．nov．林氏刀锹（新种） 450

Dorcus lhoba group 452

116．Dorcus lhoba Huang ＆ Chen sp．nov．珞巴大锹（新种） 452

117．Dorcus menba Huang ＆ Chen sp．nov．门巴大锹（新种） 455

Dorcus zhangmuensis group 456

118．Dorcus zhangmuensis Huang ＆ Chen sp．nov．樟木大锹（新种） 456

119．Dorcus nosei Nagai,2000察隅大锹 457

120．Dorcus cuonaensis Huang ＆ Chen sp．nov．错那大锹（新种） 460

Dorcus reichei group 462

121．Dorcus mencius（Kriesche,1935）孟子大锹 463

122．Dorcus liyingbingi Huang ＆ Chen sp．nov．李氏大锹（新种） 467

123．Dorcus reichei Hope,1842瑞奇大锹 469

124．Dorcus cervulus（Boileau,1901）拟瑞奇大锹 485

125．Dorcus hirticornis（Jakowlew,［1897］）毛角大锹 487

Dorcus hirticornis hirticornis（Jakowlew,［1897］）毛角大锹指名亚种 487

Dorcus hirticornis clypeatus Benesh,1950 stat．nov．毛角大锹台湾亚种 488

Dorcus hirticornis ssp．毛角大锹云南亚种 491

Species delimitation and diagnosis for species 492

Dorcus hansteini（Albers,1889）----not found in China 507

126．Dorcus kyawi Nagai ＆ Maeda,2009久氏大锹 509

Dorcus tityus group 511

127．Dorcus lineatopunctatus（Hope 1831）尼泊尔大锹 511

Dorcus lineatopunctatus gaoligongshanus subsp．nov．尼泊尔大锹东部亚种（新亚种） 520

Basic information of males 521

128．Dorcus tityus Hope,1842提提大锹（别名：提提乌斯大锹） 527

129．Dorcus daedalion（Didier ＆ Séguy,1953）华南大锹 531

Dorcus submolais（Hoper,1845）----not found in China 536

Dorcus fuliginosus（Didier,1928）----not found in China 538

Dorcus hyperion group 539

130．Dorcus laevidorsis Fairmaire,1888滇越大锹 539

131．Dorcus hyperion Boileau,1899海波力昂大锹 544

132．Dorcus hansi Schenk 2008基齿大锹（别名：三牙大锹） 548

133．Dorcus miwai Benesh,1936平头大锹 550

Unreliable records of Dorcini species for Chinese fauna in references 551

References cited in all chapters of this book 553

Plates 569

Index to species-group names,genus-group names and family-group names 709