Furthur research on the genera Lucanus Scopoli and Eolucanus Kurosawa from China,with the description of two new species 1
An updated checklist of Chinese species of the genera Lucanus and Eolucanus 1
Notes on some species of the genera Lucanus and Eolucanus 3
Ⅰ．Lucanus wuyishanensis Schenk武夷深山 3
Ⅱ．Lucanus liuyei Huang ＆ Chen刘氏深山 4
Ⅲ．Lucanus swinhoei Parry姬深山 5
Ⅳ．Lucanus prossi Zilioli普氏深山 5
Ⅴ．Lucanus brivioi Zilioli布氏深山 6
Ⅵ．Lucanus nobilis Didier雅深山 6
Ⅶ．Lucanus dirki Schenk大理深山 6
Ⅷ．Lucanus formosus Didier橙深山 7
Ⅸ．Lucanus fujitai Katsura ＆ Giang北越深山 7
Ⅹ．Lucanus sp．incert．墨脱未定名深山 8
Ⅺ．Lucanus delavayi Fairmaire黄鞘深山 8
Ⅻ．Lucanus parryi parryi Boileau帕瑞深山指名亚种 9
ⅩⅢ．Lucanus fujianensis Schenk微齿姬深山 9
ⅩⅣ．Lucanus fairmairei Planet四川深山 10
ⅩⅤ．Lucanus laminifer Waterhouse拉叉深山 10
ⅩⅥ．Lucanus angusticornis inclinatus Schenk弯叉深山中国亚种 11
ⅩⅦ．Lucanus hermani De Lisle巨叉深山 11
ⅩⅧ．Lucanus planeti planeti Planet普叉深山指名亚种 11
ⅩⅨ．Lucanus planeti dayaoshanensis Schenk普叉深山广西亚种 11
ⅩⅩ．Lucanus dybowski lhasaensis Schenk斑股深山四川亚种 12
ⅩⅪ．Lucanus ludivinae Boucher路氏深山 13
ⅩⅫ．Lucanus lunifer Westwood藏南深山 14
ⅩⅩⅢ．Lucanus thibetanus furcifer Arrow藏深山云南亚种 15
ⅩⅩⅣ．Lucanus thibetanus singularis Planet藏深山怒江亚种 16
ⅩⅩⅤ．Lucanus thibetanus isaki Nagai藏深山独龙江亚种 22
ⅩⅩⅥ．Lucanus thibetanus ssp藏深山片马未定亚种 23
ⅩⅩⅦ．Lucanus langi Huang,He ＆ Shi郎氏深山 23
ⅩⅩⅧ．Lucanus fryi Boileau弗瑞深山 25
ⅩⅩⅨ．Lucanus atratus Hope阿深山 25
ⅩⅩⅩ．Lucanus mearesii Hope金属深山 26
ⅩⅩⅪ．Lucanus nyshiwini bretschneideri Schenk双齿深山墨脱亚种 26
ⅩⅩⅫ．Eolucanus gracilis（Albers）短颚拟深山 27
ⅩⅩⅩⅢ．Eolucanus lesnei（Planet）尖齿拟深山 27
ⅩⅩⅩⅣ．Eolucanus mingyiae（Huang）圆齿拟深山 27
ⅩⅩⅩⅤ．Eolucanus pani（Huang）潘氏拟深山 30
ⅩⅩⅩⅥ．Eolucanus oberthuri（Planet）欧氏拟深山 30
Further notes 30
New descriptions 30
ⅩⅩⅩⅦ．Lucanus choui Huang ＆ Chen sp．nov．周氏深山（新种） 30
ⅩⅩⅩⅧ．Lucanus hewenjiae Huang ＆ Chen sp．nov．何氏深山（新种） 33
Descriptions of some new taxa of Lucanidae from China 35
A．Prismognathus zhangi Huang ＆ Chen sp．nov．张氏鬼锹（新种） 35
B．Ceruchus yingqii Huang ＆ Chen sp．nov．浅纹铠锹（新种） 36
C．Himaloaesalus Huang ＆ Chen gen．nov．喜马拉雅斑锹属（新属） 39
Phylogeny and systematics of Lucaninae （sensu Holloway,1960 ＆ 1968）from China 41
Introduction 41
Abbreviations 41
Choice of ingroup and outgroup taxa 41
Source of data and initial names 43
Morphology 43
Characters coding and data sets 45
Softwares 46
Analyses and searching results 47
Monophyla and higher taxa 62
1） Genera 62
2） Tribes 64
3） Species groups in Lucanus 65
4） Species groups in Neolucanus 66
5） Species groups in Prosopocoilus,Dorcus and Falcicornis 66
6） Clade of Sinodendron 66
Systematics and annotated list of higher taxa of Chinese Lucanidae,with some extension to species from outside China,and with emphasis on the tribe Dorcini 66
Unsorted genera from China 66
Subfamily AESALINAE MacLeay,1819 67
Tribe Aesalini MacLeay,1819 67
Subfamily SYNDESINAE MacLeay,1819 67
Tribe Sinodendrini Mulsant,1842 67
Tribe Ceruchini Jacquelin du Val ＆ Fainnaire,1859 67
Subfamily LUCANINAE Latreille,1804 67
Tribe Platycerini Mulsant,1842 67
Tribe Noseolucanini Huang ＆ Chen,2013,new tribe 67
Tribe Aegini Huang ＆ Chen,2013,new tribe 68
Tribe Odontolabini Parry,1870 69
Tribe Figulini Burmeister,1847 69
Tribe Capreolucanini Huang ＆ Chen,2013,new tribe 69
Tribe Cyclommatini Huang ＆ Chen,2013,new tribe 70
Tribe Lucanini Latreille,1804 71
Tribe Dorcini Parry,1864 72
Genus Gnaphaloryx Burmeister,1847 72
Genus Kirchnerius Schenk,2009 72
Genus Metallactulus Ritsema,1885 73
Genus Epidorcus Seguy,1954 73
Genus Serrognathus Motschulsky,1861 74
Genus Hexarthrius Hope,1842 75
Genus Rhaetus Parry,1864 75
Genus Rhaetulus Westwood,1871 76
Genus Pseudorhaetus Planet,1899 76
Genus Prosopocoilus Hope ＆ Westwood,1845 76
Genus Miwanus Huang ＆ Chen,new genus 77
Genus Falcicornis Planet,1894 78
Genus Sinodorcus Huang ＆ Chen,new genus 78
Genus Aulacostethus Waterhouse,1869 79
Genus Dorcus MacLeay,1819 79
Discussion on some tribes from outside China 80
Key to tribes of Lucaninae from China 82
Appendix 1-List of characters used for phylogenetic analyses 84
Appendix 2-Data matrix 101
A review of Dorcini（sensu Huang ＆ Chen,2013）from China 115
Introduction 115
Key to genera of Dorcini males 115
Key to genera of Dorcini females 117
Genus Gnaphaloryx Burmeister,1847黑锈锹属 119
1．Gnaphaloryx opacus Burmeister,1847黑锈锹 119
Genus Kirchnerius Schenk,2009钳锹属 121
Key to Chinese species of Kirchnerius males 121
Key to Chinese species of Kirchnerius females 122
Kirchnerius spencei group 122
2．Kirchnerius spencei（Hope,1840）comb．nov．钳锹 122
Kirchnerius boreli group 125
3．Kirchnerius guangxii Schenk,2009 广西钳锹 125
4．Kirchnerius yangi（Fukinuki,2004）comb．nov．杨氏钳锹 127
5．Kirchnerius simianshanus（Huang ＆ Chen,2011）comb．nov．四面山钳锹 128
6．Kirchnerius boreli（Boileau,1904）comb．nov．印度钳锹 129
Genus Metallactulus Ritsema,1885姬扁锹属 131
7．Metallactulus parvulus（Hope,1845）姬扁锹 131
Genus Epidorcus Séguy,1954前锹属 132
Key to Chinese species of Epidorcus males 132
Key to Chinese species of Epidorcus females 133
8．Epidorcus gracilis（Saunders,1854）狭长前锹 134
9．Epidorcus piceipennis（Westwood,1855）拟狭长前锹 137
10．Epidorcus denticulatus（Boileau,1901）,comb．nov．单基齿前锹 139
11．Epidorcus rondoni（Bomans ＆ Lacroix,1970）,comb．nov．短基齿前锹 144
12．Epidorcus tonkinensis（Pouillaude,1913）,comb．nov．并基齿前锹 145
13．Epidorcus bidentatus（Bomans,1978）,comb．nov．双基齿前锹 147
Genus Serrognathus Motschulsky,1861扁锹属 147
Key to Chinese taxa of Serrognathus males 147
Key to Chinese taxa of Serrognathus females 148
14．Serrognathus titanus（Boisduval,1835）扁锹 149
Serrognathus titanus platymelus（Saunders,1854）扁锹典型亚种 149
Serrognathus titanus fafner（Kriesche,［1921］）扁锹越南亚种 154
Serrognathus titanus typhoniformis（Nagel,1924）扁锹云贵亚种 155
Serrognathus titanus sika（Kriesche,［1921］）扁锹台湾亚种 156
Serrognathus titanus castanicolor Motschulsky,1861扁锹东北亚种 157
Serrognathus titanus ssp．incert．1扁锹版纳亚种 159
Serrognathus titanus ssp．incert．2扁锹墨脱亚种 160
Species delimitation 161
Diagnosis for species 163
15．Serrognathus consentaneus（Albers,1886）尖腹扁锹（别名：细齿扁） 165
16．Serrognathus kyanrauensis（Miwa,1934）深山扁锹 168
Genus Hexarthrius Hope,1842六节锹属 169
Key to Chinese taxa of Hexarthrius males 169
Key to Chinese taxa of Hexarthrius females 170
17．Hexarthrius aduncus Jordan,1894印度六节锹 170
18．Hexarthrius vitalisi Didier,1925维氏六节锹 176
Hexarthrius vitalisi vitalisi Didier,1925维氏六节锹指名亚种 176
Hexarthrius vitalisi tsukamotoi Nagai,1998维氏六节锹桂北亚种 177
Hexarthrius vitalisi ssp．incert．1维氏六节锹桂西亚种 178
Hexarthrius vitalisi ssp．incert．2维氏六节锹高黎贡山亚种 178
Species delimitation 179
Subspecies 179
19．Hexarthrius partyi Hope,1842橘背六节锹（别名：橘背叉角、帕瑞六节） 179
20．Hexarthrius forsteri（Hope,1840）佛氏六节锹 182
21．Hexarthrius melchioritis Séguy,1954缅甸六节锹 183
Genus Rhaetus Parry,1864巨鹿锹属 185
22．Rhaetus westwoodi（Parry,1862）巨鹿锹（别名：金刚鹿） 185
Genus Rhaetulus Westwood,1871鹿角锹属 187
23．Rhaetulus crenatus Westwood,1871鹿角锹 187
Rhaetulus crenatus crenatus Westwood,1871鹿角锹指名亚种 187
Rhaetulus cretatus rubrifemoratus Nagai,2000鹿角锹华南亚种 189
Rhaetulus crenatus boileaui Didier,1925？鹿角锹版纳亚种 190
Rhaetulus crenatus ssp．incert．鹿角锹中缅亚种 191
Species delimitation 191
Subspecies 191
Genus Pseudorhaetus Planet,1899拟鹿锹属 192
Key to Chinese taxa of Pseudorhaetus 192
24．Pseudorhaetus sinicus（Boileau,1899）拟鹿锹（别名：漆黑鹿角） 192
Pseudorhaetus sinicus sinicus（Boileau,1899）拟鹿锹指名亚种 192
Pseudorhaetus sinicus concolor Benesh,1960拟鹿锹台湾亚种 193
Species delimitation 194
Diagnosis for species 194
25．Pseudorhaetus oberthuri Planet,1899红背拟鹿锹 194
Genus Prosopocoilus Hope,1845锯锹属 195
Key to Chinese species of Prosopocoilus males 195
Key to Chinese species of Prosopocoilus females 197
Prosopocoilus buddha group 199
26．Prosopocoilus approximatus（Parry,1864）伪布达锯锹 199
27．Prosopocoilus thibeticus（Westwood,1855）滇东南锯锹 203
28．Prosopocoilus porrectus Bomans,1978歧齿锯锹 206
29．Prosopocoilus fulgens（Didier,1927）毛刷锯锹 206
30．Prosopocoilus confucius（Hope,1842）孔夫子锯锹 208
31．Prosopocoilus giraffa giraffa（Olivier,1789）长颈鹿锯锹指名亚种 209
32．Prosopocoilus politus（Parry,1862）亮锯锹 210
33．Prosopocoilus forficula（Thomson,1856）圆翅锯锹 212
Prosopocoilus astacoides group 214
34．Prosopocoilus astacoides（Hope,1840）两点锯锹 214
Species delimitation and subspecies delimitation 214
Prosopocoilus astacoides astacoides（Hope,1840）两点锯锹指名亚种 217
Prosopocoilus astacoides castaneus（Hope,1845）两点锯锹樟木亚种 219
Prosopocoilus astacoides fraternus（Hope,1845）两点锯锹滇南亚种 221
Prosopocoilus astacoides kachinensis Bomans ＆ Miyashita,1997两点锯锹藏东南亚种 223
Prosopocoilus astacoides blanchardi（Parry,1873）两点锯锹普通亚种 224
Prosopocoilus astacoides dubernardi（Planet,1899）两点锯锹滇西北亚种 226
Other subspecies 227
Diagnosis for species 227
Prosopocoilus impressus（Waterhouse,1869）-Valid species,not found in China 229
Prosopocoilus laterinus（Didier,1928）-Not found in China 230
35．Prosopocoilus reni Huang ＆ Chen,2011任氏锯锹 231
36．Prosopocoilus yazakii Nagai,2005三色锯锹 231
Prosopocoilus inclinatus group 232
37．Prosopocoilus inclinatus inclinatus（Motschulsky,［1858］）东北锯锹 232
38．Prosopocoilus motschulskyi（Waterhouse,1869）高砂锯锹 234
39．Prosopocoilus oweni melli Kriesche,1922欧文锯锹中国亚种 236
40．Prosopocoilus doris Kriesche,［1921］平齿锯锹 238
41．Prosopocoilus suturalis（Olivier,1789）丫纹锯锹 240
42．Prosopocoilus biplagiatus（Westwood,1855）黄纹锯锹 241
43．Prosopocoilus spineus（Didier,1927）红背锯锹 243
Prosopocoilus maclellandi group 245
44．Prosopocoilus jenkinsi（Westwood 1848）毛胫四点锯锹 245
45．Prosopocoilus maclellandi（Hope,1842）四点锯锹 249
46．Prosopocoilus laterotarsus（Houlbert,1915）砂缘锯锹 251
Genus Miwanus Huang ＆ Chen,2013 gen．nov．双钩锹属（新属） 254
47．Miwanus formosanus（Miwa,1929）comb．nov．双钩锹 254
Miwanus formosanus formosanus（Miwa,1929）双钩锹指名亚种（台湾亚种） 254
Miwanus formosanus capricornus（Didier,1931）双钩锹越南亚种 255
Miwanus formosanus kishidai（Fujita,2010）双钩锹广东亚种 256
Miwanus formosanus ssp．incert．双钩锹华中未定名亚种 256
Species delimitation 257
Genus Falcicornis Planet,1894小刀锹属 257
Key to Chinese species of Falcicornis males 257
Key to Chinese species of Falcicornis females 259
48．Falcicornis neolucanoides sp．nov．圆翅小刀锹（新种） 261
Falcicornis elegantulus group 262
49．Falcicornis pseudaxis（Didier,1926）comb．nov．南红小刀锹 262
50．Falcicornis mellianus（Kriesche,［1921］）comb．nov．黄毛小刀锹 264
51．Falcicornis seguyi（De Lisle,1955）comb．nov．叉齿小刀锹 266
Falcicornis vernicatus group 269
52．Falcicornis humilis（Arrow 1935）comb．nov．西藏小刀锹 269
53．Falcicornis vernicatus（Arrow,1938）comb．nov．戟小刀锹 276
54．Falcicornis itoi（Bomans,1993）comb．nov．越南小刀锹 279
55．Falcicornis taibaishanensis（Schenk,2008）comb．nov．拟戟小刀锹 279
56．Falcicornis heyangi Huang ＆ Chen sp．nov．何洋小刀锹（新种） 280
Falcicornis bisignatus group 281
57．Falcicornis tenuecostatus（Fairmaire,1888）comb．nov．皮氏小刀锹 281
Falcicornis tenuecostatus tenuecostatus（Fairmaire,1888）皮氏小刀锹华北亚种 281
Falcicornis tenuecostatus pieli（Didier ＆ Séguy,1953）stat．nov．皮氏小刀锹华南亚种 283
Falcicornis tenuecostatus mochizukii（Miwa,1937）stat．nov．皮氏小刀锹台湾亚种（望月锹） 286
Falcicornis tenuecostatus ssp．incert．皮氏小刀锹西康亚种 286
Species delimitation and diagnosis for species 287
58．Falcicornis moellenkampi（Nagel,1924）comb．nov．莫氏小刀锹 290
Falcicornis moellenkampi moellenkampi（Nagel,1924）莫氏小刀锹指名亚种 290
Falcicornis moellenkampi bomansi（Lacroix,1981）stat．nov．莫氏小刀锹滇西亚种 291
Diagnosis for species 293
Records 294
59．Falcicornis ruficrus（De Lisle,1970）红河小刀锹 294
60．Falcicornis bisignatus（Parry,1862）束胸小刀锹 295
61．Falcicornis fulvonotatus（Parry,1862）斑胸小刀锹 300
62．Falcicornis rufonotatus （Pouillaude,1913）滇越小刀锹 302
63．Falcicornis songianus（Didier ＆ Séguy,1953）华南小刀锹 304
64．Falcicornis himalayae（Schenk 2009）宽胸小刀锹 307
Genus Sinodorcus Huang ＆ Chen,2013 gen．nov．多刺刀锹属（新属） 308
65．Sinodorcus sawaii（Tsukawaki,1999）comb．nov．多刺刀锹 308
Genus Aulacostethus Waterhouse,1869阿锹属 309
Key to species of Aulacostethus 309
66．Aulacostethus archeri Waterhouse,1869印度阿锹 310
67．Aulacostethus ruditemporalis（Houlbert,1914）四川阿锹 311
68．Aulacostethus doani Baba,2000越南阿锹 312
69．Aulacostethus tianmuxing Huang ＆ Chen sp．nov．华东阿锹（新种） 313
Genus Dorcus MacLeay,1819刀锹属、大锹属 314
Key to Chinese species of Dorcus males 315
Key to Chinese species of Dorcus females 326
Dorcus antaeus group 333
70．Dorcus schenklingi（M？llenkamp,1913）长角大锹 333
71．Dorcus antaeus Hope,1842安达大锹（别名：安达佑实大锹） 335
72．Dorcus vicinus Saunders,1854华东大锹（别名：天章大锹） 339
73．Dorcus rectus（Motschulsky,［1858］）单齿刀锹（别名：日本小锹、直牙锹） 341
Dorcus montivagus group 344
74．Dorcus davidis（Fairmaire,1887）大卫大锹 344
Dorcus rubrofemoratus group 348
75．Dorcusyamadai（MIWA,1937）台湾刀锹（别名：刀锹） 348
76．Dorcus rubrofemoratus（van Vollenhoven,1865）红腿刀锹 351
Dorcus rubrofemoratus rubrofemoratus（van Vollenhoven,1865）红腿刀锹指名亚种 351
Dorcus rubrofemoratus chenpengi（Li,1992）红腿刀锹北部亚种 352
Dorcus rubrofemoratus miyamai Nagai ＆ Okazaki,2005红腿刀锹西部亚种 353
Species delimitation and diagnosis for species 354
77．Dorcus haitschunus（Didier ＆ Séguy,1952）长齿刀锹 354
78．Dorcus arrowi（Boileau,1911）红背刀锹 356
Dorcus arrowi arrowi（Boileau,1911）红背刀锹指名亚种（滇西亚种） 356
Dorcus arrowi magdaleinae（Lacroix,1972）stat．nov．红背刀锹滇南亚种 358
Dorcus arrowi ssp．incert．红背刀锹滇东南亚种 360
Diagnosis for species 362
79．Dorcus derelictus Parry,1862短颚刀锹 363
Dorcus nepalensis group 364
80．Dorcus sinensis（Boileau,1899）中华刀锹 364
81．Dorcus semenowi Jakowlew,1900谢苗诺夫刀锹 371
82．Dorcus kikunoae Hosoguchi,2004错那刀锹 377
83．Dorcus wui Huang ＆ Chen sp．nov．吴氏刀锹（新种） 378
84．Dorcus ratiocinativus Westwood,1871弯颚刀锹 380
85．Dorcus wardi Arrow,1943沃德刀锹 381
86．Dorcus macleayii（Hope,1845）麦氏刀锹 383
87．Dorcus branaungi Nagai,2000布氏刀锹（别名：布朗刀） 386
88．Dorcus sp．incert．1二郎山刀锹 387
89．Dorcus sp．incert．2巨刀锹 388
90．Dorcus nepalensis（Hope,1831）尼泊尔刀锹 389
91．Dorcus donckieri（Boileau,1898）东氏刀锹 392
Dorcus curvidens group 393
92．Dorcus curvidens（Hope,1840）弯角大锹 393
93．Dorcus hopei（Saunders,1854）中华大锹（别名：中国大锹） 397
Dorcus hopei hopei（Saunders,1854）中华大锹指名亚种 397
Dorcus hopei formosanus Miwa,1929 stat．nov．中华大锹台湾亚种 400
Diagnosis for species 401
Subspecies 402
94．Dorcus yaksha Gravely,1915细角大锹 402
Dorcus yaksha yaksha Gravely,1915细角大锹指名亚种 402
Dorcus yaksha gracilicornis Benesh,1950 stat．nov．细角大锹东部亚种 404
Dorcus katsurai group 407
95．Dorcus katsurai Ikeda,2000扩胫大锹 408
96．Dorcus sukkiti Fukinuki,2004缅甸扩胫大锹 411
97．Dorcus sp．incert．3肘子大锹 411
98．Dorcus myinti Nagai ＆ Maeda,2009敏氏刀锹 411
Dorcus velutinus group 413
99．Dorcus velutinus Thomson,1862锈刀锹 413
100．Dorcus tenuihirsutus Kim ＆ Kim,2010北方锈刀锹 418
101．Dorcus ursulus Arrow 1938扩胸锈刀锹 419
102．Dorcus taiwanicus Nakane ＆ Makino,1985台湾锈刀锹 420
103．Docus carinulatus Nagel,1941直颚锈刀锹 422
104．Dorcus ursulae（Schenk,1996）平齿刀锹 424
105．Dorcus tanakai Nagai,2002田中氏刀锹 426
Dorcus elegans group 427
106．Dorcus elegans（Parry,1862）樟木怪刀锹 427
107．Docus motuoensis Huang ＆ Chen sp．nov．墨脱怪刀锹（新种） 429
108．Dorcus chucheni Huang ＆ Chen sp．nov．初氏怪刀锹（新种） 433
109．Dorcus wemckeni（Schenk,2008）短颚怪刀锹 434
Dorcus striatipennis group（sensu stricto） 435
110．Dorcus striatipennis（Motschulsky,1861）双齿刀锹 435
Dorcus striatipennis yushiroi Sakaino,1997双齿刀锹台湾亚种 435
Dorcus striatipennis davidi（Séguy,1954）stat．nov．双齿刀锹大陆亚种 436
Delimitation of species 439
Subspecies 441
111．Dorcus lvbu Huang ＆ Chen sp．nov．吕布刀锹（新种） 442
112．Dorcus bashanus Huang ＆ Chen sp．nov．巴山刀锹（新种） 445
113．Dorcus gongshanus Huang ＆ Chen sp．nov．贡山刀锹（新种） 446
114．Dorcus intricatus（Lacroix,1981）雅安刀锹 448
115．Dorcus linwenhsini Huang ＆ Chen sp．nov．林氏刀锹（新种） 450
Dorcus lhoba group 452
116．Dorcus lhoba Huang ＆ Chen sp．nov．珞巴大锹（新种） 452
117．Dorcus menba Huang ＆ Chen sp．nov．门巴大锹（新种） 455
Dorcus zhangmuensis group 456
118．Dorcus zhangmuensis Huang ＆ Chen sp．nov．樟木大锹（新种） 456
119．Dorcus nosei Nagai,2000察隅大锹 457
120．Dorcus cuonaensis Huang ＆ Chen sp．nov．错那大锹（新种） 460
Dorcus reichei group 462
121．Dorcus mencius（Kriesche,1935）孟子大锹 463
122．Dorcus liyingbingi Huang ＆ Chen sp．nov．李氏大锹（新种） 467
123．Dorcus reichei Hope,1842瑞奇大锹 469
124．Dorcus cervulus（Boileau,1901）拟瑞奇大锹 485
125．Dorcus hirticornis（Jakowlew,［1897］）毛角大锹 487
Dorcus hirticornis hirticornis（Jakowlew,［1897］）毛角大锹指名亚种 487
Dorcus hirticornis clypeatus Benesh,1950 stat．nov．毛角大锹台湾亚种 488
Dorcus hirticornis ssp．毛角大锹云南亚种 491
Species delimitation and diagnosis for species 492
Dorcus hansteini（Albers,1889）----not found in China 507
126．Dorcus kyawi Nagai ＆ Maeda,2009久氏大锹 509
Dorcus tityus group 511
127．Dorcus lineatopunctatus（Hope 1831）尼泊尔大锹 511
Dorcus lineatopunctatus gaoligongshanus subsp．nov．尼泊尔大锹东部亚种（新亚种） 520
Basic information of males 521
128．Dorcus tityus Hope,1842提提大锹（别名：提提乌斯大锹） 527
129．Dorcus daedalion（Didier ＆ Séguy,1953）华南大锹 531
Dorcus submolais（Hoper,1845）----not found in China 536
Dorcus fuliginosus（Didier,1928）----not found in China 538
Dorcus hyperion group 539
130．Dorcus laevidorsis Fairmaire,1888滇越大锹 539
131．Dorcus hyperion Boileau,1899海波力昂大锹 544
132．Dorcus hansi Schenk 2008基齿大锹（别名：三牙大锹） 548
133．Dorcus miwai Benesh,1936平头大锹 550
Unreliable records of Dorcini species for Chinese fauna in references 551
References cited in all chapters of this book 553
Plates 569
Index to species-group names,genus-group names and family-group names 709